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Title Music - Ron Grainer Visual Effects - Peter Logan The Doctor and a newly-regenerated Romana arrive on Skaro and discover that the Daleks are using explosive charges and a group of humanoid slave workers to mine the planet in search of their creator, Davros.
A stalemate has arisen in an interplanetary war that the Daleks are waging against the robotic Movellans, and their hope is that Davros will be able to give them the edge.
He notices that K9 is, unusually for a robot, coughing. He establishes that K9 has a form of laryngitis which is, as the Doctor points out, pointless, as a robot would have no use for such an affliction. He calls for Romana, and is surprised when Princess Astra from the preceding adventure emerges in full regal regalia.
However this is not Astra at all: The Doctor is not impressed and tries to dissuade her from "going around wearing copies of bodies". He urges Romana to try another body. She agrees and walks out of sight to do just that. Unhappy with the height, she is told by the Doctor to "lengthen it", and she leaves to try again. When she returns, she resembles a drag queen; the Doctor possibly slightly disturbed by this gives a polite, "No thank you, not today.
He advises her to wear something more sensible and stylish, and she returns in an outfit resembling his own. He is delighted, until he realises that she again resembles Astra, only in different clothing. He gives up, realising that her mind is made up, and agrees to let her resemble Astra.
The TARDIS has landed on a very rocky planet which has breathable air and hospitable conditions, but has dangerously high levels of radioactivity. The Doctor gives Romana tablets to combat the radiation and a beeper-like device to inform her when she must take her pills.
They witness what appear to be ragged-looking natives burying one of their dead. On closer inspection, they find that this deceased fellow is not what they thought: How he came to be on this rocky planet is a mystery. They see a spaceship land, and find that it has half-buried itself into the ground in a valley. Just as the Doctor and Romana are about to investigate, underground explosions force them towards the ruins. While they explore, a column falls upon the Doctor. It is too heavy for Romana to lift alone.
She agrees to reassemble the literally "brainless" K9 and get him to assist in removing the debris. Unbeknownst to her, she is being followed. Realising that she cannot reach K9, Romana turns back. Meanwhile, the Doctor is quite happily reading a book Origins of the Universe by Oolon Colluphid , and remarks that he needs to remind Romana to take her anti-radiation pills.
A troupe of silver-haired humanoids appear and point their weapons at him. He attempts to charm them, but it does not seem to work. Romana returns to the ruin.
The Doctor has vanished. As she turns to leave, she finds the man who has been following her blocking her path. She backs away, only to fall down a rubble chute, losing consciousness.
The man prepares to climb down and help her, but before he can, she recovers consciousness and hears a drilling noise coming from one of the walls. The wall seems to be moving. She backs away from the wall. Suddenly a pair of Daleks burst through it: You are our prisoner — do not move.
You are our prisoner! The man, meanwhile, has seen the whole thing. Meanwhile, the Doctor is thanking the Movellans as these silver-haired humanoids are called for helping him, and remarks at their strength. He asks their commander, Sharrel, what planet they are on, and is told that it is known as DGamma-Z-Alpha. The Doctor enquires to its name, and is astonished to hear that the answer is Skaro. The Movellans are here to wage war against the Daleks.
Romana, meanwhile, is being interrogated by the Daleks. After determining that she is no threat to them, the Daleks command Romana to work at one of their drilling sites. The Doctor and the Movellans meet with the man who has been following Romana and him. He identifies himself as Starship Engineer Tyssan, captured by the Daleks two years ago.
He collapses after revealing that the Daleks have used him as slave labour for drilling as part of a search operation. He soon comes around, and says he does not know what the Daleks are looking for.
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He tells the Doctor about what has happened to Romana, and they set out to rescue her. She learns that she is getting weaker as a result of radiation sickness, and is told that the only way out of captivity is to die. Within minutes, she collapses and seems to die. Her fellow workers remove her body.
As the Doctor frantically tries to dig her out, Romana appears and explains that she feigned death in order to escape. At school she had been taught how to stop her hearts. They head to the Dalek headquarters.
Lan is left on guard outside of the Control Centre, and is shot down by a Dalek who is out searching for them. The Doctor establishes that the Daleks are searching for something on a level that they have yet to access. He remembers an alternative route to this area, so he, Romana, and Agella make their way to this floor while Sharrel returns to his ship. They discover Davros, the creator of the Daleks, who had seemingly been exterminated when the Doctor last saw him.
Genesis of the Daleks. Something gives way up above, and part of the ceiling collapses on Agella.
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While the party is distracted by this, Davros starts to stir: Episode Three "The resurrection has come, as I always knew it would", says the awakened Davros. The Doctor finds Davros and takes him into a blocked-off room in the old Dalek city. He lets Romana and Tyssan escape out the window, and they return to the Movellan ship. The Daleks find them and proceed to exterminate their prisoners until the Doctor complies. The Doctor threatens to kill Davros with a makeshift explosive he has just concocted.
He orders the Daleks to free all their prisoners, and to let him escape. The Daleks say that these conditions are unacceptable and illogical - and therefore, to a Dalek, impossible. The exterminations will continue. The Daleks now comply. Davros frantically orders the Daleks to remove the explosive, which they do. The Doctor detonates the explosive remotely, and the explosion seems to take a Dalek with it. Davros vows to make the Daleks invincible, and the supreme power of the Universe. Unbeknownst to them, Agella is not dead.
She returns and reports all she has just heard to her fellow Movellans. Romana reaches the Movellan spaceship, but learns that the Movellans are not as altruistic as they appear. Agella uses her weapon on her and knocks her out. The Doctor meets up with Tyssan and they find a Movellan scout. He finds that the unconscious Romana has been attached to the nova device, sealed inside an airtight container. He sends Tyssan away and tries to open the container, as the timer is ticking down However, the nova device was revealed to be a "dud" — a decoy used to lure the Doctor.
The Daleks want Davros to help them gain an advantage. The Movellans want the Doctor to do the same for them, which the Doctor refuses to do. Davros, on the other hand, is eager to give the Daleks the upper hand; he orders them to make a suicide bombing attack on the Movellan craft on realising that the Doctor might do the same thing for the Movellans.
The Doctor leads an attack by the slaves on the Movellans, which ends with them all being deactivated. While the prisoners take control of the Movellan ship, the Doctor makes his way to the city to confront Davros. He tells Davros that the Movellans have been disabled; unfortunately Davros does not believe him and intends to destroy the Movellan ship anyway. The slaves are no match for the Daleks, who begin exterminating them.
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As the Dalek fires around blindly nearly killing Davros in the process , the Doctor attaches an explosive to it and blows it up, then activates the bomb detonator and destroys the attacking Dalek squad. He takes Davros into the custody of the former slave workers. Davros shall be placed in Cryogenic suspension and taken to Earth to stand trial for his crimes.
The Doctor and Romana leave, remarking on the fact that whoever makes mistakes often wins as the Doctor knows only too well.